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Ph.D. Courses and Curriculum

Program Requirements

  • Two core courses: QCB 515 and COS/QCB 551
  • Three additional courses from the lists below, including at least one each from the Quantitative and Biological course lists 
  • QCB Graduate Colloquium
  • Responsible Conduct of Research (RCR) summer course: QCB 501
  • Research rotations (a minimum of two required)
  • Oral general exam
  • At least two semesters of teaching
  • Completion of thesis (progress overseen by thesis committee)
The course of study for each student must be approved by the Director of Graduate Studies in the beginning of their first year. Course substitutions are possible with the permission of the DGS.
Quantitative Courses (must take at least one)  
  • CHM 515 Biophysical Chemistry I
  • CHM/MOL 550 Contemporary Problems in Molecular Biophysics
  • COS 511 Theoretical Machine Learning
  • COS 557 Analysis & Visualization of Large-Scale Genomic Datasets
  • MOL 510 Introduction to Biological Dynamics
  • MOL 536 Advanced Statistics for Biology
  • PHY 561/2 Biophysics
  • QCB/CBE 511 Modeling Tools for Cell and Developmental Biology
Biological Courses (must take at least one) 
  • CHM/QCB 541 Chemical Biology II
  • EEB 504 Fundamental Concepts in Ecology, Evolution, and Behavior II
  • EEB 507 Recent Research in Population Biology
  • MOL 504 Cellular Biochemistry
  • MOL 505 Molecular Biology of Prokaryotes
  • MOL 506 Molecular Biology of Eukaryotes
  • MOL 516 Genetics of Eukaryotic Organisms
  • MOL 520 Advanced Topics in Prokaryotic and Eukaryotic Cell Biology
  • MOL 523 Molecular Basis of Cancer  
  • MOL 545 Advanced Microbial Genetics
  • MOL 559 Viruses: Strategy & Tactics 
Selected undergraduate courses of interest
(Note: these do not count towards course requirements)

For additional information regarding courses, please visit the Office of the Registrar's Course Offerings page.