Wurzburger, N., Bellenger, J.P, Kraepiel, A.M.L. and L.O. Hedin. 2012. Molybdenum and phosphorus interact to constrain asymbiotic nitrogen fixation in tropical forests. PLoS ONE 7(3): e33710. doi:10.1371/journal.pone.0033710 (pdf)

Brookshire, E.N.J., Hedin, L.O., Newbold, J.D., Sigman, D.M. and J.K Jackson. 2012. Sustained losses of bioavailable nitrogen from montane tropical forests.  Nature Geoscience 5, 123-126. (pdf)

Bernal, S., Hedin, L.O., Likens, G.E., Gerber, S. and D.C. Buso. 2012. Complex response of the forest nitrogen cycle to climate change.  Proceedings of the National Academy of Sciences 109 (9): 3406-11. (pdf)

Brookshire, E.N.J., Gerber, S., Menge D.N.L. and L.O. Hedin. 2011. Large losses of inorganic nitrogen from tropical rainforests suggest a lack of nitrogen limitation.  Ecology Letters doi: 10.1111/j.1461-0248.2011.01701. (pdf)

Wright, S.J., Yavitt., J.B., Wurzburger, N., Turner, B.L., Tanner, E.V.J., Sayer, E.J., Santiago, L.W., Kaspari, M., Hedin, L.O., Harms, K.E., Garcia, M.N. and M.D. Corre. 2011. Potassium, phosphorus or nitrogen limit root allocation, tree growth and litter production in a lowland tropical forest. Ecology 92, 1616-1625. (pdf)

Barron, A.R., Purves, D.W. and L.O. Hedin. 2011. Facultative nitrogen fixation by canopy legumes in a lowland tropical forest.  Oecologia, 165, 511-520. (pdf)

Gerber, S., Hedin, L.O., Oppenheimer, M., Pacala, S.W. and E. Shevliakova. 2010. Nitrogen cycling and feedbacks in a global dynamic land model. Global Biogeochemical Cycles 24, GB1001, doi 10.1029/2008GB003336.  (pdf)

Reich, P.B., J. Oleksyn, I.J. Wright, K.J. Niklas, L. Hedin, and J.J. Elser. 2010. Evidence of a general 2/3-power leaf nitrogen to phosphorus scaling among major plant groups and biomes. Proc. Roy. Soc. Ser. B. 277: 877-883. (pdf)

Gerber, S., L.O. Hedin, M. Oppenheimer, S.W. Pacala, E. Shevliakova. 2010 Nitrogen cycling and feedbacks in a global dynamic land model. Global Biogeochemical Cycles, 24, GB1001, doi:10.1029/2008GB003336. (pdf)

Hedin, L.O., E.N.J. Brookshire, D.N.L. Menge, and A.R. Barron. 2009. The nitrogen paradox in tropical forests ecosystems. Annu. Rev. Ecol. Evol. Syst.. 40:613–35. (pdf)

Menge, D.N.L.,S.A. Levin, & L.O. Hedin. 2009. Facultative versus obligate nitrogen fixation strategies and their ecosystem consequences. American Naturalist 174: 465-477. Epub doi:10.1086/605377. (pdf)

Menge, D.N. L. and L.O. Hedin. 2009. Nitrogen fixation in different biogeochemical niches along a 120,000-year chronosequence in New Zealand. Ecology  90 (8):  2190-2201. Epub doi: 10.1890/08-0877.1 (pdf)

Menge, D.N.L., S.W. Pacala and L.O. Hedin. 2009.  Emergence and Maintenance of Nutrient Limitation over Multiple Timescales in Terrestrial Ecosystems.  American Naturalist  173 (2): 164-175. (pdf)

Barron, A.R., N. Wurzburger, J-P Bellenger, S.J. Wright, A.M. Kraepiel, and L.O. Hedin. 2009. Molybdenum limitation of asymbiotic nitrogen fixation in tropical forest soils.  Nature Geoscience. 2:42-45. Epub doi:10.1038/ngeo366. [Article was also published as the journal cover article in the January 2009 print version of Nature Geoscience]. (pdf)

Menge, D.N., S.A. Levin, and L.O. Hedin. 2008. Evolutionary tradeoffs can select against nitrogen fixation and thereby maintain nitrogen limitation. Proceedings of the Natural Academy of Sciences 105 (5):1573-8. (pdf)

Perring, M.P., L.O. Hedin, S.A. Levin, M. McGroddy and C. de Mazancourt. 2008. Increased plant growth from nitrogen addition should conserve phosphorus in terrestrial ecosystems. Proceedings of the Natural Academy of Science. 105:1971-1976 (pdf)

McGroddy, M.E., W.T. Baisden and L.O. Hedin. 2008. Stoichiometry and hydrological C, N, and P losses across climate and geology: an environmental matrix approach across New Zealand primary forests. Global Biogeochemical Cycles, 22: GB1026, doi:10.1029/2007GB003005. (pdf)

Houlton, B, Sigman, D.M, Schuur, E.A.G. and Hedin, L.O. 2007. A climate-driven switch in plant nitrogen acquisition within tropical forest communities. Proceedings of the National Academy of Sciences 104:8902-8906. (pdf)

Perakis, S.S. and L.O. Hedin. 2007. State factor relationship of dissolved organic carbon and nitrogen losses from unpolluted temperate forest watersheds.  Journal of Geophysical Research- Biogeochemisstry 112:G02010 APR 25 2007. (pdf)

Hedin, L.O. 2007. Does the exception prove the rule? Reply.  Nature 445: E11. (pdf)

von Fischer, J.C. and Hedin, L.O. 2007. Controls on soil methane fluxes: tests of biophysical mechanisms using stable isotope tracers. Global Biogeochemical Cycles 21: GB2007 MAY 4 2007. (pdf)

Hedin, L.O. 2006. Physiology: Plants on a different scale. Nature 439:399-400. (pdf)

Houlton, B., Sigman, D. and Hedin, L.O. 2006. Isotopic evidence for large gaseous nitrogen losses from tropical rainforests. Proceedings of the National Academy of Sciences 103: 8745-8750. (pdf)

Houlton, B, Sigman, D, Schuur, T. and Hedin, L.O. 2007. A climate-driven switch in plant nitrogen acquisition within tropical forest communities. Proceedings of the National Academy of Sciences. (pdf)

Daufresne, T. and Hedin, L.O. 2005. Plant coexistence depends on ecosystem nutrient cycles: extension ofthe resource-ratio theory. Proceedings of the National Academy of Sciences 102: 9212-9217. (pdf)

Perakis, S.S., Compton, J.E., Hedin, L.O. 2005. Nitrogen retention across a gradient of N-15 additions to an unpolluted temperate forest soil in Chile. Ecology, 86: 96-105. (pdf)

Hedin, L.O. 2004. Global organization of terrestrial plant-nutrient interactions. Proceedings of the National Academy of Sciences 101:10849-10850. (Invited commentary) 11 (pdf)

Jackson, R.B. and Hedin, L.O. 2004. Terrestrial and Freshwater Biogeochemistry. Ecology, 9:2353-2354. (Special feature) (pdf)

McGroddy, M.E., Daufresne, T., and Hedin, L.O. 2004. Scaling of C:N:P: stoichiometry in forest ecosystems worldwide: implications of terrestrial Redfield-type ratios. Ecology, 85: 2390-2401. (Special feature) (pdf)

Hedin, L.O., Vitousek, P.M. and Matson, P.A. 2003. Nutrient losses over four million years of tropical forest development. Ecology 84(9):2231-2255. (pdf)

Perakis, S.S., and Hedin, L.O. 2002. Nitrogen loss from unpolluted South American forests mainly via dissolved organic compounds. Nature 415:416-419. (pdf)

Perakis, S.S., and Hedin, L.O. 2002. Response to commentary on “Nitrogen loss from unpolluted South American forests mainly via dissolved organic compounds.” Nature 418:665-665. (pdf) Cover: (pdf)

Kennedy, M.J., Hedin, L.O. and Derry, L.A. 2002. Decoupling of unpolluted temperate forests from rock nutrient sources revealed by natural 87Sr/86Sr and 84Sr tracer addition. Proceedings of the National Academy of Sciences 99: 9639-9644.(pdf)

von Fischer, J.C. and Hedin, L.O. 2002. Separating methane production and consumption with a fieldbased isotope pool dilution technique. Global Biogeochemical Cycles 16: 8-1 to 8-13. (pdf)

Ostrom, N.E., Hedin, L.O., von Fischer, J.C. and Robertson, G.P. 2002. Nitrogen transformations and NO3- removal at a soil-stream interface: A stable isotope approach. Ecological Applications 12: 1027-1043. (pdf)

Hedin, L.O., Hobbie, S.E., Jackson, R.B. and Neff, J.C. 2002. What role will Ecology (and the ESA) play in the future of Earth Sciences? The Bulletin of the Ecological Society of America 83:201-202. (pdf)

Neff, J. and Hedin, L.O. 2002. Building a home for the biogeosciences: Challenges for understanding terrestrial ecosystems. EOS 83:165-166. (pdf)

Hedin, L.O., Chadwick, O., Schimel, J. and Torn, M. 2002. Linking Ecological Biology and Geoscience: Challenges for terrestrial environmental science. Princeton Report to the National Science Foundation: 16 pp. (pdf)

Perakis, S.S. and Hedin, L.O. 2001. Fluxes and Fates of Nitrogen in Soil of an Unpolluted Old-Growth Temperate Forest, southern Chile. Ecology 82:2245-2260. (pdf)

Salmon, C.D., Hedin, L.O., Walter, M.T. and Brown, M.G. 2001. Hydrological controls on chemical export from an undisturbed old-growth Chilean forest. Journal of Hydrology 253:69-80. (pdf)

Hedin, L.O. 2000. Deposition of nutrients and pollutants to ecosystems. In Methods in Ecosystem Science, Sala, O.E., Jackson, R.B., Mooney, H.A. and Howarth, R.W., eds., pp. 265-276 Springer Verlag, New York, N.Y. (pdf)

Chadwick, O.A., Derry, L.A., Vitousek, P.M., Huebert, B.J. and Hedin, L.O. 1999. Changing sources of nutrients during four million years of ecosystem development. Nature 397:491-497. (pdf)

Cleveland, C.C., Townsend, A.R., Schimel, D.S., Fisher, H., Howarth, R.W., Hedin, L.O., Perakis, S.S., Latty, E.F., von Fischer, J.C., Elseroad, A. and Wasson, M.F. 1999. Global patterns of terrestrial biological nitrogen (N2) fixation in natural ecosystems. Global Biogeochemical Cycles 13:623-645. (pdf)

Walter, M.T., Mutch, P., Salmon, C.D., McCool, D.K. and Hedin, L.O. 1999. Digitizing chart recorder data: coordinate system conversion for rain gauges and similar recording instruments. Journal of Atmospheric and Oceanic Technology 16:1138-1142. pdf

Hedin, L.O., von Fischer, J.C., Ostrom, N.E., Kennedy, B.P., Brown, M.G. and Robertson, G.P. 1998. Thermodynamic constraints on nitrogen transformations and other biogeochemical processes at soilstream interfaces. Ecology 79:684-703. pdf

Hedin, L.O. 1998. From landscapes to the sea: Challenges to understanding how humans influence aquatic ecosystems. In Proceedings of the 7th Stockholm Water Symposium and 3rd International Conference on the Environmental Management of Enclosed Coastal Seas, Stockholm, Sweden, Falkenmark, M.ed., pp. 33-44. The Stockholm International Water Institute and Arkpressen. pdf

Ostrom, N.E., Knoke, K.E., Hedin, L.O., Robertson, G.P. and Smucker, A.J.M. 1998. Temporal trends in nitrogen isotope values of nitrate leaching from an agricultural soil. Chemical Geology 146:219-227. pdf

Perez, C.A., Hedin, L.O. and Armesto, J.J. 1998 Nitrogen mineralization in two unpolluted old-growth forests of contrasting biodiversity and dynamics. Ecosystems 1:361-373. pdf

Sobczak, W.V., Hedin, L.O. and Klug, M.J. 1998. Relationships between bacterial productivity and organic carbon at a soil-stream interface. Hydrobiologia 386:45-53. pdf

Vitousek, P.M., Hedin, L.O., Matson, P.A., Fownes, J.H. and Neff, J. 1998. Within-system element cycles, input-output budgets, and nutrient limitation. In Successes, Limitations, and Frontiers in Ecosystem Science, Pace, M. L. and Groffmann, P.M. eds., pp. 432-451. Springer-Verlag, New York.

Findlay, S., Likens, G.E., Hedin, L.O., Fisher, S.G. and McDowell, W.H. 1997. Organic matter dynamics in Bear Brook, Hubbard Brook Experimental Forest, New Hampshire, USA. Journal of the North American Benthological Society 16:43-46. pdf

Hedin, L.O. and Likens, G.E. 1996. Atmospheric dust and acid rain. Scientific American 275:88-92. pdf

Hedin, L.O. and Hetherington, E. 1996. Atmospheric and geologic constraints on the biogeochemistry of North and South American temperate rain forests. In High-Latitude Rainforests and Associated Ecosystems of the West Coast of the Americas, Lawford, R.G., Alaback, P.B. and Fuentes, E., eds., pp. 57-74. Springer-Verlag, New York. pdf

Hedin, L.O., Armesto, J.J. and Johnson, A.H., 1995. Patterns of nutrient loss from unpolluted, old-growth temperate forests: Evaluation of biogeochemical theory. Ecology 76:493-509. pdf

Armesto, J.J., Villagrán, C., Aravena, J.C., Pérez, C., Smith-Ramirez, C., Cortés, M. and Hedin, L.O. 1995. Conifer forests of the Chilean coastal range. In Ecology of the Southern Conifers, Enright, N.J. and Hill, R.S. eds., pp. 156-170. Melbourne University Press, Melbourne, Australia

Hedin, L.O. 1994. Stable isotopes, unstable forest. Nature 372:725-726. pdf

Hedin, L.O., Granat, L., Likens, G.E., Buishand, T.A., Galloway, J.N., Butler, T.J., and Rodhe, H. 1994. Steep declines in atmospheric base cations in regions of Europe and North America. Nature 367:351-354. pdf

Hedin, L.O. and Brown, M.B. 1994. Watershed-level coupling of nitrogen and sulfur in a heterogeneous landscape. Verhandlungen der International Vereinungen Limnologie 25:1477-1482.

Hedin, L.O. and Campos, H. 1991. Importance of small streams in understanding and comparing watershed ecosystem processes. Revista Chilena de Historia Natural 64:583-596.

Hedin, L.O. 1991. Cross-system comparisons of detritus food webs. In Comparative Analyses of Ecosystems: Patterns, Mechanisms and Theories, Cole, J.J., Lovett, G.M. and Findlay, S. eds., pp. 346-347. Springer Verlag, New York.

Hedin, L.O., Likens, G.E., Postek, K.M. and Driscoll, C.T. 1990. A field experiment to test whether organic acids buffer acid deposition. Nature 345:798-800. pdf

Hedin, L.O., Granat, L., Likens, G.E. and Rodhe, H. 1990. Strong similarities in seasonal concentration ratios of SO4 2-, NO3 -, and NH4 + in precipitation between Sweden and the northeastern United States. Tellus 42B:454-462. pdf

Likens, G.E., Bormann, F.H., Hedin, L.O., Driscoll, C.T. and Eaton, J.S. 1990. Dry deposition of sulfur: A 23-year record for the Hubbard Brook Forest ecosystem. Tellus 42B:319-329. pdf

Hedin, L.O. 1990. Factors controlling sediment community respiration in woodland stream ecosystems. Oikos 57:94-105. pdf

Likens, G.E., Hedin, L.O. and Butler, T.J. 1990. Some long-term precipitation chemistry patterns at the Hubbard Brook Experimental Forest: Extremes and averages. Verhandlungen der International Vereinungen Limnologie 24:128-135.

Fox, T. D., Folley, L.S., Mulero, J.J., McMullin, T.W., Thorsness, P.E., Hedin, L.O. and Costanzo, M.C. 1990. Analysis and manipulation of yeast mitochondrial genes. Methods in Enzymology 194:149-165. pdf

Driscoll, C.T., Likens, G.E., Hedin, L.O., Eaton, J.S., and Bormann, F.H. 1989. Changes in the chemistry of surface waters. Environmental Science and Technology 23:137-143. pdf

Driscoll, C.T., Likens, G.E., Hedin, L.O. and Bormann, F.H. 1989. Response to letter by Chen and Gomez. Environmental Science and Technology 23:754-789. pdf

Driscoll, C.T., Likens, G.E., Hedin, L.O. and Bormann, F.H. 1989. Response to letter by Dillon.
Environmental Science and Technology 23:1028. pdf

Driscoll, C.T., Likens, G.E., Hedin, L.O. and Bormann, F.H. 1989. Response to letter by Holdren and Church. Environmental Science and Technology 23:1079-1080.

Hedin, L.O., Mayer, M.S. and Likens, G.E. 1988. The effect of deforestation on organic debris dams. Verhandlungen der International Vereinungen Limnologie 23:1135-1141.

Hedin, L.O., Likens, G.E. and Bormann, F.H. 1987. Decrease in precipitation acidity resulting from decreased SO4 2- concentration. Nature 325:244-246 pdf