An introduction to the concepts, approaches, and methods for studying complex ecological systems from local to global scales. Students will examine nutrient cycling, energy flow, and evolutionary processes, with emphasis on experimental approaches and comparisons between terrestrial, freshwater, and marine ecosystems. Particular attention will be on effects of human activities including climate change, biodiversity loss, eutrophication, and acid rain. Lectures cover theoretical elements and examples from the primary literature. Laboratories offer hands-on exposure to state-of-the-art field methods, and include an independent project.