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Department of Ecology & Evolutionary Biology
Levin Lab Mailing Address: 106A Guyot Hall
Princeton, NJ 08544-1003
Phone: (609) 258-6879. Fax (609) 258-6819


EEB logo
Front, L-R: Caroline Farrior (Ph.D. 2012), Eili Klein (Ph.D. 2012), Allison Shaw (Ph.D. 2012), and Liliana Salvador (Ph.D. 2012). Back, L-R: Simon Leblanc (graduate student), Simon A. Levin, Carla Staver (Ph.D. 2012).
Note: EEB = Dept. of Ecology & Evolutionary Biology; PACM = Program in Applied & Computational Mathematics


Wohl, Philip R. (1971)
(joint advisor with Sol Rubinow)
Applied Mathematics

Thesis title: The Traverse Force on a Drop in Unbounded Poiseuille Flow
Deceased 1996

Udovic, J. Daniel (1973)
(joint advisor with D. Pimentel)

Thesis title: Evolution in Predator-Prey Systems: Some Extensions of the Genetic Feedback Model
Current position: Professor Emeritus, Department of Biology and Environmental Studies, University of Oregon

Sastre, Antonio (1974)
Applied Mathematics

Thesis title: Analysis and Simulation of Pacemaker Neurons
Current position: Scientific Review Officer, Center for Scientic Review, NIH, HHS

Hastings, Alan M. (1977)
Applied Mathematics

Thesis title: Some Models in Population Biology
Current position: Professor; Chair, Division of Environmental Studies, University of California, Davis

Gross, Louis, J. (1979)
Applied Mathematics

Thesis title: Models of the Photosynthetic Dynamics of Fragaria virginiana
Current position: Professor, Department of Mathematics, University of Tennessee

Runkle, James, R. (1979)

Thesis title: Gap Phase Dynamics in Climax Mesic Forests
Current position: Professor, Department of Biology, Wright State University

White III, George N. (1980)
Applied Mathematics

Thesis title: A Mathematical Study of the Role of Chemotaxis in an Intertidal Predator-Prey System
Current position: Biomathematician, Bedford Institute of Oceanography

Kareiva, Peter M. (1981)
(joint advisor with Richard Root)

Thesis title: Non-Migratory Movement and the Distribution of Herbivorous Insects: Experiments with Plant Spacing and the Application of Diffusion Models to Mark-Recapture Data
Current position: Chief Scientist; Director of Science, The Nature Conservancy

Nedelman, Jerry R. (1981)
(joint advisor with Sol Rubinow)
Applied Mathematics

Thesis title: I. Examination of the Kinetic Support of the Two-State Model of the Cell Cycle II. Facilitated Diffusion of Oxygen
Current position: Director, Novartis Pharmaceuticals

Ellner, Stephen P. (1981)
Applied Mathematics

Thesis title: Evolutionary Stable Germination Behaviors in Randomly Varying Environments
Current position: Professor, Dept. of Ecology and Evolutionary Biology, Cornell University

Castro Ospina, Jose Mildred (1984)
Applied Mathematics

Thesis title: Equilibrium and Stability of Bioeconomic Models of Renewable Resources Under Diverse Harvesting Regimes
Current: Retired

Craig, Catherine L. (1985)

Thesis title: Predator-Prey Dynamics: The Role of Spider and Insect Ecological and Behavioral Interactions in the Evolution of the Araneoidea
Current position: Research Associate, Museum of Comparative Zoology, Harvard University; President, Conservation Through Poverty Alleviation, International

Liu, Wei-min (1987)
Applied Mathematics

Thesis title: Dynamics of Epidemiological Models-Recurrent Outbreaks in Autonomous Systems
Current position:Director; Biostatistician, Roche Molecular Systems, Inc. (RMI)

Andreasen, Viggo A. (1988)
Applied Mathematics
Thesis title: Dynamical Models of Epidemics in Age-Structured Populations-Analysis and Simplification
Current position: Professor, Department of Science, Roskilde University, Denmark

Braner, Moshe (1988)
Thesis title: Dormancy, Dispersal and Staged Development: Ecological and Evolutionary Aspects of Structured Population in Random Environments
Current position: Statistical Analyst, Public Health Surveillance, Vermont Department of Health

Cain, Michael L. (1989)

Thesis title: Patterns of Clonal Growth in Solidago altissima and Medeola virginiana
Current position: Research Associate in Biology and Mathematics, Bowdoin College

Adler, Frederick R. (1991)
Applied Mathematics
Thesis title: Models of Structured Populations
Current position: Professor, Departments of Mathematics and Biology, University Utah

Nuernberger, Beate (1991)

Thesis title: Population Structure of Dineutus assimilis in a Patchy Environment: Dispersal, Gene Flow, and Persistence
Current position: Lecturer, Ludwig-Maximilians-Universität, München, Germany

Grunbaüm, Daniel (1992)
Thesis title: Local Processes and Global Patterns: Biomathematical Models of Bryzoan Feeding Currents and Density Dependent Aggregations in Antarctic Krill
Current position: Adjunct Associate Professor, Department of Biology; Associate Professor, School of Oceanography, University of Washington

Limburg, Karin E. (1994)
Thesis title: Ecological Constraints on Growth and Migration of Juvenile American Shad (Alosa sapidissima Wilson) in the Hudson River Estuary, New York
Current position: Associate Professor, Deptartment of Environmental Science and Forest Biology, SUNY College of Environmental Science and Forestry, Syracuse

Deutschman, Douglas (1996)
Thesis title: Scaling From Trees to Forests: The Problem of Relevant Detail
Current position: Professor, Department of Biology,
San Diego State University

Grevstad, Fritzi S. (1998)
(joint advisor with Richard Root)
Thesis title: The Colonization Ecology of Two Loosestrife Leaf Beetles, Galerucella calmariensis and G. pusilla
Current position: Research Scientist, Olympic Natural Resources Center, University of Washington


Gandhi, Amar (1996)
(co-advisor with Stephen Orszag)
Thesis title: From Level to Level: Course-Graining/Scaling in Ecology
Current position: Group Project Manager, Google, Inc.

Palsson, Eirikur (1996)
(co-advisor with Edward Cox)

Thesis title: The cAMP Signaling System in Dictyostelium discoideum
Current position: Assistant Professor, Deptartment of Biology, Simon Fraser University

Dushoff, Jonathan (1997)

Thesis title: Modeling the Effects of Host Heterogeneity on the Spread of Human Diseases
Current position: Assistant Professor, Department of Biology, McMaster University

Smith, David L. (1998)

Thesis title: The Evolutionary Ecology of Parasitic Plants
Current position: Visiting Professor, Epidemiology, Oxford University

Solorzano, Luis A. (1999)
Thesis title: Analyzing Vegetation-Environment Associations from Regional to Local Scales: A Case Study in Eastern Amazonia (Brazil)
Current position: Program Director, Environmental Science, Moore Foundation

Muller-Landau, Helene (2001)

Thesis title: Seed Dispersal in a Tropical Forest: Empirical Patterns, Their Origins, and Their Consequences for Community Dynamics
Current position: Staff Scientist, Smithsonian Tropical Research Institute

Ma, Junling (2002)

Thesis title: Evolutionary Branching and Its Application to Resource Adaptation
Current position: Assistant Professor, Departments of Mathematics and Statistics, University of Victoria, Canada

Malvadkar, Urmila (2002)

Thesis title: Variation in Diel Vertical Migration of Zooplankton: Causes and Consequences
Current position: Assistant Professor, St. Olaf College, MN

Chan, Kai M.A. (2003)
Thesis title: The Effects of Slightly Leaky Prezygotic Isolating Barriers and the Use of Phylogenetic Tree Shape to Study Diversification
Current position: Canada Research Chair; Assistant Professor, Institute for Resources, Environment and Sustainability, University British Columbia

Cline, Jon C. (2003)
Thesis title: Community Structure and Patch Dynamics in Rocky Intertidal Communities: Linking Process and Pattern Across Scales
Current position: Information Systems Engineer, Lead, MITRE, Center for Connected Government, VA

Keymer, Juan E. (2003)

Thesis title: Competition for Space and Intertidal Community Structure
Current position: Investigator, Centro de Estudios Cientificos Caldivia, Chile

Plotkin, Joshua (2003)
Thesis title: Aggregation in Ecology and Evolutionary Biology
Current position: Associate Professor, University of Pennsylvania, Departments of Biology and Computer Science

Worden, Lee (2003)
Thesis title: Evolution, Constraint, Cooperation, and Community Structure in Simple Models
Current position: Postdoctoral Researcher, MacMaster University and UC Berkley; Visiting Professor, San Francisco Art Institute

Zea-Cabrera, Eduardo (2003)
Thesis title: Plant Water Use Strategies: Physiological Mechanisms and Ecological Dynamics
Current position: Director, Humboldt Research Institute, Columbia

Jolles, Anna E. (2004)
(co-advisor with Andy Dobson)
Thesis title: Disease Ecology of Tuberculosis in African Buffalo
Current position: Assistant Professor, College of Veterinary Medicine, Oregon State University

Leslie, Nandi (2005)
Thesis title: Spatial Stochastic Models for Landscape Degradation and Deforestation in Bolivia and Brazil
Current position: Operations Research Analyst, Systems Planning and Defense, Inc., Washington, DC

Baskett, Marissa (2006)

Thesis title: Marine Reserve Design and Life History Variation
Current position: Assistant Professor, Department of Environmental Science and Policy, University of California, Davis

Livnat, Adi (2006)
Thesis title: Causes and Consequences of Internal Conflict: A Theoretical Behavioral Perspective
Current position: Assistant Professor, VPI

Bazykin, Georgii (2007)

Thesis title: Natural Selection in Molecular Evolution: Genome Scale, Parsimony-Based Approach
Current position: Research Fellow, Institute for Information Transmission Problems, Russian Academy of Sciences, Moscow, Russia

Jeremy Lichstein (2007)
(co-advisor with Steve Pacala)

Thesis title: Theoretical and Empirical Perspectives on the Dynamics of U.S. Forests
Current position: Assistant Professor, University of Florida, Gainsville, Department of Biology

Juliet Pulliam (2007)
Thesis title: Determinants and Dynamics of Viral Host Jumps
Current position:Assistant Professor, University of Florida, Gainsville, Department of Biology

Benjamin Strauss (2007)
Thesis title: Snails at Three Scales: Interplay of Stream Hydrology and Hydraulics with the Morphology, Dispersal and Distribution of Elimia proxima
Current position: Associate Director, Strategic Initiatives, Climate Control, Princeton, NJ

Kryazhimskiy, Sergey (2008)
Thesis title: Pathogen Evolution under Natural Selection: The Influenza A Case Study
Current position: Postdoctoral Researcher, Harvard University

Menge, Duncan (2008)
(co-advisor with Lars Hedin)

Thesis title: The Ecology and Evolution of Nitrogen Fixation and Nutrient Limitation at Multiple Time Scales
Current position: Assistant Professor, Dept. of Ecology, Evolution and Environmental Biology, Columbia University

Ndifon, Wilfred (2009)

Thesis title: Quantitative Approaches to Influenza Virus Surveillance
Current position: Career Development Chair, Quantitative Immunology, African Institute for Mathematical Sciences (AIMS), South Africa and AIMS-Ghana

Chisholm, Ryan (2010)

Thesis title: The Ecology and Economics of Alien Invasive Species with a Case Study from the South African Fynbos
Current position: Assistant Professor, National University of Singapore

de Froment, Adrian (2010)
Thesis title: Fighting for Information: Decision-Making, Animal Contests and the Emergence of Social Hierarchy
Current position: Barrister, Serle Court, London, England

Nadell, Carey (2011)
Thesis title: The Evolution of Social Behavior in Bacterial Biofilms
Current position: Howard Hughes Medical Insitute Postdoctoral Scientist, Princeton University, Department of Molecular Biology (PI: Bonnie L. Bassler); Visiting Research Fellow (Isaac Newton Institute, Cambridge University, UK (Fall 2014)

Staver, Ann Carla (2012)
(co-advisor with Lars Hedin)


Thesis title: Determinants and Dynamics of the Global Distribution of the Savanna and Forest Biomes
Current position: Assistant Professor, Dept. of Ecology and Evolutionary Biology, Yale University

Klein, Eili Y. (2012)
Thesis title: Antimalarial Drug Resistance
Current position: Assistant Professor, Center for Advanced Modeling in the Social, Behavioral and Health Sciences, Dept. of Emergency Medicine, Johns Hopkins University

Shaw, Allison (2012)
(co-advisor with Iain Couzin)
Thesis title: Modeling Motives for Movement: Theory for Why Animals Migrate
Current position: Assistant Professor, Dept. of Ecology, Evolution and Behavior, University of Minnesota

Farrior, Caroline (2012)
(co-advisor with Steve Pacala)
Thesis title: The Roles of Competition and Resource Availability in the Biomass Allocation of Plant Communities
Current position: Postdoctoral Researcher, National Institute for Mathematical and Biological Synthesis (NIMBioS), Knoxville, TN

Salvador, Liliana (2012)
(co-advisor with Francisco Dionisio)
Thesis title: Searching Without Information: A Quantitative Analysis of Caenorhabditis elegans
Current position: Postdoctoral Researcher, University of Glasgow, Scotland

Berdahl, Andrew (2014)
(co-advisor with Iain Couzin)
Thesis title: Collective Navigation
Current position: Omidyar Fellow, Santa Fe Institute, Santa Fe, NM


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