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submitted manuscripts


  1. [PDF] Clonal dominance in excitable cell networks
    J. Imran Alsous, J. Rozman, R. Marmion, A. Košmrlj, and S. Y. Shvartsman, Nature Physics, in press
  2. [PDF] Method of image charges for describing deformation of bounded two-dimensional solids with circular inclusions
    S. Sarkar, M. Čebron, M. Brojan, and A. Košmrlj, Physical Review E 103, 053004 (2021)
  3. [PDF] Elastic multipole method for describing deformation of infinite two-dimensional solids with circular inclusions
    S. Sarkar, M. Čebron, M. Brojan, and A. Košmrlj, Physical Review E 103, 053003 (2021)
  4. [PDF] Local accumulation of extracellular matrix regulates global morphogenetic patterning in the developing mammary gland
    B. A. Nerger, J. M. Jaslove, H. E. Elashal, S. Mao, A. Košmrlj, A. J. Link, and C. M. Nelson, Current Biology 31, 1-15 (2021)
  5. [PDF] Buckling of thermalized elastic sheets
    A. Morshedifard, M. Ruiz-Garcia, M. J. Abdolhosseini Qomi, and A. Košmrlj, Journal of the Mechanics and Physics of Solids 149, 104296 (2021)


  1. [PDF] Designing the Morphology of Separated Phases in Multicomponent Liquid Mixtures
    S. Mao, M. S. Chakraverti-Wuerthwein, H. Gaudio, and A. Košmrlj, Physical Review Letters 125, 218003 (2020) [News]
  2. [PDF] Size-dependent patterns of cell proliferation and migration in freely-expanding epithelia
    M. A. Heinrich, R. Alert, J. M. LaChance, T. J. Zajdel, A. Košmrlj, and D. J. Cohen, eLife 9, e58945 (2020)
  3. [PDF] Finite Temperature Phase Behavior of Viral Capsids as Oriented Particle Shells
    A. Singh, A. Košmrlj, and R. Bruinsma, Physical Review Letters 124, 158101 (2020)
  4. [PDF] Non-uniform growth and surface friction determine bacterial biofilm morphology on soft substrates
    C. Fei, S. Mao, J. Yan, R. Alert, H.A. Stone, B.L. Bassler, N.S. Wingreen, and A. Košmrlj, Proceedings of the National Academy of Sciences of the United States of America 117 (14), 7622-7632 (2020) [News]


  1. [PDF] Smooth muscle differentiation shapes domain branches during mouse lung development
    K. Goodwin, S. Mao, T. Guyomar, E. Miller, D. C. Radisky, A. Košmrlj, and C. M. Nelson, Development 146, dev181172 (2019)
  2. [PDF] Mechanical Instability and Interfacial Energy Jointly Drive Biofilm Morphogenesis
    J. Yan, C. Fei, S. Mao, A. Moreau, N. S. Wingreen, A. Košmrlj, H. A. Stone, and B. L. Bassler, eLife 8, e43920 (2019)
  3. [PDF] Phase behavior and morphology of multicomponent liquid mixtures
    S. Mao, D. Kuldinow, M. P. Haataja, and A. Košmrlj, Soft Matter 15, 1297-1311 (2019)


  1. [PDF] Heterogeneous Absorption of Antimicrobial Peptide LL37 in Escherichia coli Cells Enhances Population Survivability
    M. Snoussi, J. P. Talledo, N.-A. Del Rosario, S. Mohammadi, B.-Y. Ha, A. Košmrlj, and S. Taheri-Araghi, eLife 7, e38174 (2018) [News1], [News2], [News3]
  2. [PDF] Bacterial Biofilm Materials Properties Enable Removal and Transfer by Capillary Peeling
    J. Yan, A. Moreau, S. Khodaparast, A. Perazzo, J. Feng, C. Fei, S. Mao, A. Košmrlj, N. S. Wingreen, B. L. Bassler, and H. A. Stone, Advanced Materials 30, 1804153 (2018) [News]
  3. [PDF] Enhanced cellular uptake of LHRH-conjugated PEG-coated magnetite nanoparticles for specific targeting of triple negative breast cancer cells
    J. Hu, J. D. Obayemi, K. Malatesta, A. Košmrlj, and W. O. Soboyejo, Materials Science and Engineering C 88, 32-45 (2018)


  1. [PDF] Spontaneous formation of aligned DNA nanowires by capillarity-induced skin folding
    S. Nagashima, H.D. Ha, D.H. Kim, A. Košmrlj, H.A. Stone, and M.-W. Moon, Proceedings of the National Academy of Sciences of the United States of America 114, 6233-6237 (2017)
  2. [PDF] Non-Hookean statistical mechanics of clamped graphene ribbons
    M. J. Bowick, A. Košmrlj, D. R. Nelson and R. Sknepnek, Physical Review B 95, 104109 (2017)
  3. [PDF] Statistical Mechanics of Thin Spherical Shells
    A. Košmrlj, and D. R. Nelson, Physical Review X 7, 011002 (2017)


  1. [PDF] Asymmetric Damage Segregation Constitutes an Emergent Population-Level Stress Response
    S. Vedel, H. Nunns, A. Košmrlj, S. Semsey and A. Trusina, Cell Systems 3, 187-198 (2016)
  2. [PDF] Response of thermalized ribbons to pulling and bending
    A. Košmrlj, and D. R. Nelson, Physical Review B 93, 125431 (2016)


  1. [PDF] A monomer-trimer model supports intermittent glucagon fibril growth
    A. Košmrlj, P. Cordsen, A. Kyrsting, D. E. Otzen, L. B. Oddershede, and M. H. Jensen, Scientific Reports 5, 9005 (2015)


  1. [PDF] Complex ordered patterns in mechanical instability induced geometrically frustrated triangular cellular structures
    S. H. Kang, S. Shan, A. Košmrlj, W. L. Noorduin, S. Shian, J. C. Weaver, D. R. Clarke and K. Bertoldi, Physical Review Letters 112, 098701 (2014) [News]
  2. [PDF] Thermal excitations of warped membranes
    A. Košmrlj and D. R. Nelson, Physical Review E 89, 022126 (2014)


  1. [PDF] Harnessing instabilities for design of soft reconfigurable auxetic/chiral materials
    J. Shim, S. Shan, A. Košmrlj, S. H. Kang, E. R. Chen, J. C. Weaver and K. Bertoldi, Soft Matter 9, 8198-8202 (2013)
  2. [PDF] Mechanical properties of warped membranes
    A. Košmrlj and D. R. Nelson, Physical Review E 88, 012136 (2013)
  3. [PDF] Statistical Physics of T-Cell Development and Pathogen Specificity
    A. Košmrlj, M. Kardar and A. K. Chakraborty, Annual Review of Condensed Matter Physics 4, 339-360 (2013)


  1. [PDF] The Influence of T Cell Development on Pathogen Specificity and Autoreactivity
    A. Košmrlj, M. Kardar and A. K. Chakraborty, Journal of Statistical Physics 149, 203-219 (2012)


  1. [PDF] Continuum theory for cluster morphologies of soft colloids
    A. Košmrlj, G. J. Pauschenwein, G. Kahl and P. Ziherl, Journal of Physical Chemistry B 115, 7206-7217 (2011)
  2. [PDF] Thymic selection of T cells as diffusion with intermittent traps
    A. Košmrlj, Journal of Statistical Physics 142, 1277-1286 (2011)


  1. [PDF] Effects of thymic selection of the T-cell repertoire on HLA class I-associated control of HIV infection
    A. Košmrlj, E. L. Read, Y. Qi, T. M. Allen, M. Altfeld, S. G. Deeks, F. Pereyra, M. Carrington, B. D. Walker and A. K. Chakraborty, Nature 465, 350-354 (2010)
  2. [PDF] Statistical Mechanical Concepts in Immunology
    A. K. Chakraborty and A. Košmrlj, Annual Review of Physical Chemistry 61, 283-303 (2010)


  1. [PDF] How a protein searches for its site on DNA: the mechanism of facilitated diffusion
    L. Mirny, M. Slutsky, Z. Wunderlich, A. Tafvizi, J. Leith and A. Košmrlj, Journal of Physics A: Mathematical and Theoretical 42, 434013 (2009)
  2. [PDF] Thymic Selection of T-Cell Receptors as an Extreme Value Problem
    A. Košmrlj, A. K. Chakraborty, M. Kardar and E. I. Shakhnovich, Physical Review Letters 103, 068103 (2009)


  1. [PDF] How the thymus designs antigen-specific and self-tolerant T cell receptor sequences
    A. Košmrlj, A. K. Jha, E. S. Huseby, M. Kardar and A. K. Chakraborty, Proceedings of the National Academy of Sciences of the United States of America 105, 16671-16676 (2008)


  1. [PDF] Soft spheres make more mesophases
    M. A. Glaser, G. M. Grason, R. D. Kamien, A. Košmrlj, C. D. Santangelo and P. Ziherl, Europhysics Letters 78, 46004 (2007)